Richard Etheridge (in memorium; obituary here)

Emeritus Professor of Biology
Department of Biology, San Diego State University
Evolutionary Biology Program Area
Ph.D., University of Michigan

Biography: Espinoza, R. E. and de Queiroz, K. 2008. Historical Perspectives: Richard Emmett Etheridge. Copeia 3: 708–717.

Research Interests and Activities:
Morphology, systematics, and evolution of iguanian lizards, currently focused on the families Polychrotidae and Tropiduridae
   in austral South America.


Etheridge, R.  1948.  Range of the Faded Snake, Arizona elegans elegans  in Texas. Herpetologica 4(6): 149.

Etheridge, R. 1948. The reptiles and amphibians of Aransas County, Texas. Annual Report Texas Game, Fish and Oyster Commission for 1948.         

Etheridge, R. 1950.  Color variants in snakes from southeastern United States. Copeia 1950(4): 321.

Etheridge, R., R. Anderson and E. A. Liner.  1952  Notes on amphibian and reptile populations in a Louisiana pineland area. Ecology 33(2): 274-278.

Etheridge, R.  1952. The southern range of the racer Coluber constrictor stejnegerianus (Cope), with remarks on the Guatemalan species Coluber ortenburgeri  Stuart.  Copeia 1952(3):189-190.

Etheridge, R.  1952. The Warty Gecko, Hemidactylus turcicus turcicus  (Linnaeus) in New Orleans, Louisiana. Copeia 1952(1):47-48.

Smith, H. M.  and R. Etheridge.  1953.  Resurrection of Plestiodon Belli Gray (Reptilia: Squamata: Lacertilia) for a Mexican skink.  Herpetologica. 8: 153-161

Schwartz, A., and R. Etheridge. 1954.  New and additional herpetological records from the North Carolina coastal plain. Herpetologica 10(3): 167-171.

Etheridge, R. 1958.  Pleistocene lizards of the Cragin Quarry Fauna, Meade County, Kansas. Copeia 1958(2): 94-101.

Etheridge, R.  1960.  The Slender Glass Lizard, Ophisaurus attenuatus  from the Pleistocene (Illinoian Glacial) of Oklahoma. Copeia 1960(1): 46-47.

Etheridge, R. 1960.  Additional notes on the lizards of the Cragin Quarry Fauna.  Papers of the Michigan Academy of Arts and Letters 45: 113-117.

Etheridge, R.  1960.  The Pliocene lizards genus Eumecoides Taylor.  Bulletin of the Southern California Academy of Sciences 59(2): 62-69.

Etheridge, R.  1961.  Additions to the herpetological fauna of Cerralvo Island in the Gulf of California, Mexico.  Herpetologica 17(1): 57-60.

Etheridge, R.   1961.  Late Cenozoic grass lizards (Ophisaurus ) in the southern Great Plains.  Herpetologica 17(3): 179-186.

Etheridge, R.  1962.  "Pan de Azucar" in the Peruvian rain forest.  Los Angeles County Museum Quarterly Report (1) 1: 4-7.     

Etheridge, R.  1962.  Skeletal variation in the iguanid lizard Sator grandaevus.  Copeia 1962 (3): 613-619.

Etheridge, R.  1964.  The skeletal morphology and systematics relationships of sceloporine lizards.  Copeia 1964(4): 610-631.

Etheridge, R.  1964.  Late Pleistocene lizards from Barbuda, British West Indies.  Bulletin of the Florida Academy of Sciences 9(2): 43-75.

Etheridge, R.  1965.  The abdominal skeleton of lizards in the family Iguanidae.  Hereptologica, 21(3): 161-168

Etheridge, R.  1965.  Fossil lizards of the Dominican Republic.  Quartely Journal of the Florida Academy of Sciences 28 (1): 84-105.

Etheridge, R.  1966  Pleistocene lizards from New Providence. Quartely Journal of the Florida Academy of Sciences 28 (4): 349-358.

Etheridge, R.  1966. The systematic relationships of West Indian and South American lizards referred to the iguanid genus Leiocephalus.  Copeia 1966(1): 79-91.

Etheridge, R.  1966.  An extinct lizard of the genus Leiocephalus  from Jamaica. Quartely Journal of the Florida Academy of Sciences 29(1): 47-59.

Etheridge, R.  1967.  Lizard Caudal vertebrae.  Copeia, 1967(4): 699-721.

Etheridge, R.  1968.  A review of the iguanid lizard genera Uracentron  and Strobilurus.  Bulletin of the British Museum (Natural history), Zolology, London 17 (2): 45-64. 

Etheridge, R.  1969.  Tropidogaster blainvillii  Duméril & Bibron, 1837 (Reptilia, Sauria) proposed suppression under plenary powers. Bulletin of Zoological Nomenclature, London  25(6): 224-246.

Etheridge, R.  1969.  A review of the iguanid lizard genus Enyalius. Bulletin of the British Museum (Natural history), Zolology, London 18(8): 233-260.

Etheridge, R. 1970.  A review of the South American iguanid lizard genus Plica. Bulletin of the British Museum (Natural history), Zolology, London 19(7): 235-246.

Etheridge, R.  1970.  Enyalius  (pp. 117-118), Ophryoessoides  (pp. 212-215), Plica  (pp. 230-231), Stenocercus  (pp. 254-257), Tropidurus  (pp. 265-269), Uracentron  (pp. 273-274). In Peters, J. A. and R. Donoso-Barros (eds.).  Catalogue of the Neotropical Squamata: Part II. Lizards and  Amphisbaenians, Bulletin of the United States National Museum 297:viii+293 pp.  

Etheridge, R.  1982.   A checklist of the iguanine and Malagasy iguanid lizards (pp. 5-37). In Burghardt, G. M.  and A. S. Rand (eds.) Iguanas of the world, Noyes Publications, Park Ridge, New Jersey xix+472 pp.

Cei, J. M., R. Etheridge, and F. Videla.  1983.  Especies nuevas de iguanido del noroeste de la provincia de San Juan (Reserva Provincial San Guillermo) Argentina.  Deserta 7:316-323

Etheridge, R., and E. E. Williams.  1985.  Notes on Pristidactylus (Squamata: Iguanidae).  Breviora, Museum of Comparative Zoology, Harvard, Cambridge 483: 1-18.

Etheridge, R. and K. de Queiroz.  1988.  A phylogeny of Iguanidae (pp. 285-367). In Estes. R. D. and G. Pregill (eds.) Phylogenetic relationships of the lizard families, essays commemorating Charles L. Camp, Stanford University Press, Stanford xv+631 pp.

Frost, D and R. Etheridge.  1989.  A phylogenetic analysis and taxonomy of iguanian lizards (Reptilia: Sqamata).  University of Kansas Museum of Natural history, Miscellaneous Publications 81: 65 pp.

Etheridge, R.  and E. E. Williams.  1991.  A review of the South American lizard genera Urostrophus  and Anisolepis  (Squamata: Iguania: Polychridae).  Bulletin of the Museum of Camparative Zoology 152((5): 317-361

Etheridge, R.  and E. E. Williams.  1992.   Laemanctus obtusirostris Wiegmann, 1834 and Laemanctus fitzingeri Wiegmann, 1834 (Reptilia, Squamata) proposed supression under the plenary powers. Bulletin of Zoological Nomenclature 49(3): 217-220.

Etheridge, R.  1992.  A new psammophilus lizard of the genus Liolaemus (Squamata: Tropiduridae)  from northwestern Argentina. Bollettino del Museo Regionale di Scienze Naturali, Torino 10(1): 1-19.

Frost, D. and R. Etheridge.  1992.  A consideration of iguanian lizards and the objectives of systematics: a reply to Lazell.  Herpetological Review  23(2): 50-54.

Etheridge, R.  1993.  Lizards of the Liolaemus darwinii  complex (Squamata: Tropiduridae) in northern Argentina. Bollettino del Museo Regionale di Scienze Naturali, Torino 11(1): 137-199.

Etheridge, R.  1995.  Redescription of Ctenoblepharys adspersa Tschudi 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae).  American Museum of Natural History Novitates Novitates 3142, 34 pp.

Etheridge, R., and R. E Espinoza.  1997.  New records and natural history notes for lizards from northwestern Argentina.  Herpetological Review 28(3) 160-161.

Etheridge, R.  1998.  Redescripton and status of Liolaemus hatcheri Stejneger, 1909 (Reptilia: Squamata: Tropiduridae). Cuadernos de Herpetología 12(1) 160-161.

Etheridge, R.  1998.  Redscription and status of Liolaemus hatcheri Stejneger, 1909 (Reptilia: Squamata: Tropduridae). Cuadernos de Herpetología 12(1): 31-36.

Etheridge, R.  2000.  A review of lizards of the Liolaemus wiegmannii group (Squamata, Tropiduridae), and a history of morphological change in the sand-dwelling species.  Herpetological Monographs 14:293-352.

Etheridge, R. and R. E. Espinoza.  2000.  Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semi-annotated bibliography.  Smithsonian Herpetological Information Service 127, 64 pp.

Etheridge, R.  2001. A new species of Liolaemus (Reptilia: Squamata: Tropiduridae) from Mendoza Province, Argentina. Cuadernos de Herpetología 15(1): 3-15.

Frost, D., R. Etheridge, D. Janies, and T. A. Titus.  2001.  Total evidence, sequence alignment, evolution of polychrotid lizards, and a reclassification of the Iguania (Squamata:I guania). American Museum Novitates 3343, 38 pp.

Vlladares, J. P., R. Etheridge, J. Schulte II, G. Manriquez, and A. Spotorino. 2002. Nueva especie de lagartija del norte de Chile, Liolaemus molinai (Reptilia: Liolaeminae). Revista Chilena de Historía Natural 75: 473-489.

Wiens, J. and R. Etheridge.  2003.  Phylogenetic relationships of hoplocercid lizards, coding and combining meristic, morphometric, and polymorphic data using step matices. Herpetologica 30(3): 375-398. 

Etheridge, R. and M. Christie.  2003.  Two new species of the lizard genus Liolaemus (Squamata: Liolaemidae) from northern Patagonia, with comments on Liolaemus rothi.  Journal of Herpetology 37(2): 325-341.

Etheridge, R. and J. M. Savage.  2003.  Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma: Repilia, Sauria): proposed conservtion of usage of the names by designation of a neotype for Lacerta palluma Molina, 1782.  Bulletin of Zoological Nomenclature 60(1): 38-41.

Cabrera, M. R. and R. Etheridge.  2006.  New records and type locality restriction for the endemic Argentinian lizard Cnemidophorus tergolaevigatus (Squamata: Teiidae).  Herpetological Review 37(1): 110-111.

Etheridge, R. and J. M. Savage.  2006.  Stability restored to the name Phymaturus palluma: a response to Cei and Scolaro (2006).  Herpetological Review 37(4): 406-407.

Torres-Carvajal, O., K. de Queiroz, and R. Etheridge.  2009. A new species of lizard (Iguanidae: Hoplocercinae: Enyalioides) from the upper basin of Río Zamora in southeastern Ecuador.  ZooKeys, 27: 59-71.

Espinoza, R., F. Lobo, and R. Etheridge. 2011. Taxonomic history of the iguanian lizard  Liolaemus pictus major Boulenger with a revalidation of Liolaemus capillitas Hulse.  Journal of Herpetology. 45: 129-133.

Torres-Carvajal, O.,  R. Etheridge and K. de Queroz.  2011. A systematic revision of neotropical lizards in the clade Hoplocercinae (Squamata: Iguania). Zootaxa 2752: 1-44.

Lobo, F. and R. Etheridge. 2013. The type locality of Phymaturus palluma (Molina, 1782) (Reptilia: Iguania: Liolaemidae) and the status of Phymaturus adrianae Pereyra, 1992 and other unnamed populations. South American Journal of Herpetology 8(2): 89-101.

Etheridge and Frost. 2010-11. Checklists of the Pleurodont Iguanians.
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Troncoso-Placios, J. and R. Etheridge. 2012. Distributional range of the poorly known Liolaemus tacnae (Shreve 1931). Herpetological Bulletin 121: 35-38.